1. Osmólska, Halszka, 1982, Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia: Neues Jahrbuch für Geologie und Paläontologie - Monatshefte.

BibTeX
@article{doi101127njgpm19821982440,
    author = "Osmólska, Halszka",
    title = "Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia",
    year = "1982",
    journal = "Neues Jahrbuch für Geologie und Paläontologie - Monatshefte",
    url = "https://doi.org/10.1127/njgpm/1982/1982/440",
    doi = "10.1127/njgpm/1982/1982/440",
    openalex = "W3092107547"
}

2. Osmolska, H, 1982, Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia.

BibTeX
@misc{osmolska1982hulsanpes1,
    author = "Osmolska, H",
    title = "Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia",
    year = "1982",
    howpublished = "Neues Jb. Geol. Palaeont. Mh., p. 440-448",
    note = "talkorigins\_source = {true}; raw\_reference = {Osmolska, H., 1982, Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia: Neues Jb. Geol. Palaeont. Mh., p. 440-448.}"
}

3. Osmólska, Halszka, 1982, Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia: Neues Jahrbuch für Geologie und Paläontologie - Monatshefte: v. 1982, no. 7: p. 440-448.

BibTeX
@article{osmólska1982hulsanpes,
    author = "Osmólska, Halszka",
    title = "Hulsanpes perlei n.g. n.sp. (Deinonychosauria, Saurischia, Dinosauria) from the Upper Cretaceous Barun Goyot Formation of Mongolia",
    year = "1982",
    journal = "Neues Jahrbuch für Geologie und Paläontologie - Monatshefte",
    url = "https://doi.org/10.1127/njgpm/1982/1982/440",
    doi = "10.1127/njgpm/1982/1982/440",
    number = "7",
    openalex = "W3092107547",
    pages = "440-448",
    volume = "1982"
}

4. Jenkins, Farish A. and Schaff, Charles R., 1988, The Early Cretaceous mammal Gobiconodon (Mammalia, Triconodonta) from the Cloverly Formation in Montana: Journal of Vertebrate Paleontology.

Abstract

ABSTRACT Gobiconodon ostromi, sp. nov., described from two partial skeletons collected from the Cloverly Formation in southcentral Montana, is closely related to Gobiconodon borissiaki Trofimov (1978) from the Early Cretaceous of Mongolia. The unusual dental characters of this new form are enlarged caniniform incisors, reduced canines, and replacement of the molariform teeth that lie distal to the presumptive premolars. Despite the fact that the molariform occlusal pattern is the same as in Amphilestes of the Middle Jurassic and the overall structure of the molariform teeth is comparable to that in amphilestid triconodonts, the dentition is otherwise so anomalous that Gobiconodon is assigned to a new family. The scapula possesses a supraspinous fossa and an apparently large coracoid. The humerus exhibits large deltopectoral and medial crests, and protuberant epicondyles; the planes of the proximal and distal halves of this bone are substantially offset. The pelvis conforms to a generalized mammalian pattern. A deep fossa of unknown function occurs on the dorsal surface of the lesser trochanter; the greater trochanter extends distally well down onto the shaft. A spur-like bone appears to represent an extratarsal element. In overall body size, Gobiconodon ostromi, sp. nov., is comparable to Didelphis virginiana but in its limb and vertebral proportions is more robust.

BibTeX
@article{doi10108002724634198810011681,
    author = "Jenkins, Farish A. and Schaff, Charles R.",
    title = "The Early Cretaceous mammal Gobiconodon (Mammalia, Triconodonta) from the Cloverly Formation in Montana",
    year = "1988",
    journal = "Journal of Vertebrate Paleontology",
    abstract = "ABSTRACT Gobiconodon ostromi, sp. nov., described from two partial skeletons collected from the Cloverly Formation in southcentral Montana, is closely related to Gobiconodon borissiaki Trofimov (1978) from the Early Cretaceous of Mongolia. The unusual dental characters of this new form are enlarged caniniform incisors, reduced canines, and replacement of the molariform teeth that lie distal to the presumptive premolars. Despite the fact that the molariform occlusal pattern is the same as in Amphilestes of the Middle Jurassic and the overall structure of the molariform teeth is comparable to that in amphilestid triconodonts, the dentition is otherwise so anomalous that Gobiconodon is assigned to a new family. The scapula possesses a supraspinous fossa and an apparently large coracoid. The humerus exhibits large deltopectoral and medial crests, and protuberant epicondyles; the planes of the proximal and distal halves of this bone are substantially offset. The pelvis conforms to a generalized mammalian pattern. A deep fossa of unknown function occurs on the dorsal surface of the lesser trochanter; the greater trochanter extends distally well down onto the shaft. A spur-like bone appears to represent an extratarsal element. In overall body size, Gobiconodon ostromi, sp. nov., is comparable to Didelphis virginiana but in its limb and vertebral proportions is more robust.",
    url = "https://doi.org/10.1080/02724634.1988.10011681",
    doi = "10.1080/02724634.1988.10011681",
    openalex = "W2041142773",
    references = "doi101002aja1001370304, doi101002jmor1051670308, doi101002jmor1051850203, doi101098rstb19760022, doi101111j109636421981tb01127x, doi101111j109636421985tb01500x, doi101111j1469185x1968tb00966x, doi105281zenodo16386718, doi105962bhltitle3460, openalexw1539913220"
}

5. Russell, Dale A. and Dong, Zhi-Ming, 1993, A nearly complete skeleton of a new troodontid dinosaur from the Early Cretaceous of the Ordos Basin, Inner Mongolia, People's Republic of China: Canadian Journal of Earth Sciences.

Abstract

An articulated skeleton of a 1 m long theropod from Early Cretaceous strata in Inner Mongolia is clearly referrable to the Troodontidae, representing the most complete specimen known of this group of small carnivorous dinosaurs. The tail and neck of the animal were curled next to its body upon burial; the upper surface of the skeleton was badly damaged by erosion. Previously unknown details of troodont morphology include a quadratic contact with the braincase wall (forming part of a channel leading to the lateral depression), a presacral vertebral count possibly similar to that of most theropods, absence of ossified caudal tendons, presence of a rod-like clavicle, and absence of sternal ossifications. A new genus and species (Sinornithoides youngi n.gen., n.sp.) is established for the specimen on the basis of its short skull, laterally directed orbital rim of the frontal, and elongated hind limb. A reassessment of character distributions in other small theropods and primitive birds must be completed before troodontid affinities can be established with greater precision.

BibTeX
@article{doi101139e93187,
    author = "Russell, Dale A. and Dong, Zhi-Ming",
    title = "A nearly complete skeleton of a new troodontid dinosaur from the Early Cretaceous of the Ordos Basin, Inner Mongolia, People's Republic of China",
    year = "1993",
    journal = "Canadian Journal of Earth Sciences",
    abstract = "An articulated skeleton of a 1 m long theropod from Early Cretaceous strata in Inner Mongolia is clearly referrable to the Troodontidae, representing the most complete specimen known of this group of small carnivorous dinosaurs. The tail and neck of the animal were curled next to its body upon burial; the upper surface of the skeleton was badly damaged by erosion. Previously unknown details of troodont morphology include a quadratic contact with the braincase wall (forming part of a channel leading to the lateral depression), a presacral vertebral count possibly similar to that of most theropods, absence of ossified caudal tendons, presence of a rod-like clavicle, and absence of sternal ossifications. A new genus and species (Sinornithoides youngi n.gen., n.sp.) is established for the specimen on the basis of its short skull, laterally directed orbital rim of the frontal, and elongated hind limb. A reassessment of character distributions in other small theropods and primitive birds must be completed before troodontid affinities can be established with greater precision.",
    url = "https://doi.org/10.1139/e93-187",
    doi = "10.1139/e93-187",
    openalex = "W2005056616"
}

6. Byers, Alton C., 1996, Historical and Contemporary Human Disturbance in the Upper Barun Valley, Makalu-Barun National Park and Conservation Area, East Nepal: Mountain Research and Development: v. 16, no. 3: p. 235.

BibTeX
@article{byers1996historical,
    author = "Byers, Alton C.",
    title = "Historical and Contemporary Human Disturbance in the Upper Barun Valley, Makalu-Barun National Park and Conservation Area, East Nepal",
    year = "1996",
    journal = "Mountain Research and Development",
    url = "https://doi.org/10.2307/3673946",
    doi = "10.2307/3673946",
    number = "3",
    pages = "235",
    volume = "16"
}

7. XU, XING and CHOINIERE, JONAH N. and PITTMAN, MICHAEL and TAN, QINGWEI and XIAO, DONG and LI, ZHIQUAN and TAN, LIN and CLARK, JAMES M. and NORELL, MARK A. and HONE, DAVID W. E. and SULLIVAN, CORWIN, 2010, A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China: Zootaxa: v. 2403, no. 1.

Abstract

We describe a new dromaeosaurid theropod from the Upper Cretaceous Wulansuhai Formation of Bayan Mandahu, Inner Mongolia. The new taxon, Linheraptor exquisitus gen. et sp. nov., is based on an exceptionally well-preserved, nearly complete skeleton. This specimen represents the fifth dromaeosaurid taxon recovered from the Upper Cretaceous Djadokhta Formation and its laterally equivalent strata, which include the Wulansuhai Formation, and adds to the known diversity of Late Cretaceous dromaeosaurids. Linheraptor exquisitus closely resembles the recently reported Tsaagan mangas. Uniquely among dromaeosaurids, the two taxa share a large, anteriorly located maxillary fenestra and a contact between the jugal and the squamosal that excludes the postorbital from the infratemporal fenestra. These features suggest a sister-taxon relationship between L. exquisitus and T. mangas, which indicates the presence of a unique dromaeosaurid lineage in the Late Cretaceous of Asia. A number of cranial and dental features seen in L. exquisitus and T. mangas, and particularly some postcranial features of L. exquisitus, suggest that these two taxa are probably intermediate in systematic position between known basal and derived dromaeosaurids. The discovery of Linheraptor exquisitus is thus important for understanding the evolution of some salient features seen in the derived dromaeosaurids.

BibTeX
@article{xu2010a,
    author = "XU, XING and CHOINIERE, JONAH N. and PITTMAN, MICHAEL and TAN, QINGWEI and XIAO, DONG and LI, ZHIQUAN and TAN, LIN and CLARK, JAMES M. and NORELL, MARK A. and HONE, DAVID W. E. and SULLIVAN, CORWIN",
    title = "A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China",
    year = "2010",
    journal = "Zootaxa",
    abstract = "We describe a new dromaeosaurid theropod from the Upper Cretaceous Wulansuhai Formation of Bayan Mandahu, Inner Mongolia. The new taxon, Linheraptor exquisitus gen. et sp. nov., is based on an exceptionally well-preserved, nearly complete skeleton. This specimen represents the fifth dromaeosaurid taxon recovered from the Upper Cretaceous Djadokhta Formation and its laterally equivalent strata, which include the Wulansuhai Formation, and adds to the known diversity of Late Cretaceous dromaeosaurids. Linheraptor exquisitus closely resembles the recently reported Tsaagan mangas. Uniquely among dromaeosaurids, the two taxa share a large, anteriorly located maxillary fenestra and a contact between the jugal and the squamosal that excludes the postorbital from the infratemporal fenestra. These features suggest a sister-taxon relationship between L. exquisitus and T. mangas, which indicates the presence of a unique dromaeosaurid lineage in the Late Cretaceous of Asia. A number of cranial and dental features seen in L. exquisitus and T. mangas, and particularly some postcranial features of L. exquisitus, suggest that these two taxa are probably intermediate in systematic position between known basal and derived dromaeosaurids. The discovery of Linheraptor exquisitus is thus important for understanding the evolution of some salient features seen in the derived dromaeosaurids.",
    url = "https://doi.org/10.11646/zootaxa.2403.1.1",
    doi = "10.11646/zootaxa.2403.1.1",
    number = "1",
    openalex = "W2157947199",
    volume = "2403",
    references = "doi101038nature02898, doi101038nature03996, doi101038nature08124, doi101038nature08322, doi101126science1144066, doi101126science28454232137, doi1015259780520941434, doi102475ajss319111253, doi105281zenodo16171435, doi105860choice353642"
}

8. Bhattacharya, Sabyasachi, 2014, Professor Barun De (1932–2013): Indian Historical Review: v. 41, no. 1: p. 147-150.

BibTeX
@article{bhattacharya2014professor,
    author = "Bhattacharya, Sabyasachi",
    title = "Professor Barun De (1932–2013)",
    year = "2014",
    journal = "Indian Historical Review",
    url = "https://doi.org/10.1177/0376983614521724",
    doi = "10.1177/0376983614521724",
    number = "1",
    pages = "147-150",
    volume = "41"
}

9. Cau, Andrea and Madzia, Daniel, 2018, Redescription and affinities of Hulsanpes perlei (Dinosauria, Theropoda) from the Upper Cretaceous of Mongolia: PeerJ: v. 6: p. e4868.

Abstract

Hulsanpes perlei is an enigmatic theropod dinosaur from the Baruungoyot Formation (?mid- to upper Campanian, Upper Cretaceous) of Mongolia. It was discovered in 1970, during the third Polish-Mongolian paleontological expedition to the Nemegt Basin. The taxon is known based on a partial braincase and an incomplete right hindlimb. However, the braincase fragment has never been described nor illustrated. We redescribe all elements that form the holotype of Hulsanpes and discuss the affinities of this taxon. The braincase fragment is interpreted as belonging to the inner ear region, and includes the floccular recess and part of the labyrinth. Hulsanpes perlei is confirmed as a valid taxon, diagnosed by a unique combination of metatarsal characters, including two autapomorphies. Historically, it represents the oldest record of the recently-established clade Halszkaraptorinae. Our findings identify subcursorial adaptations for Hulsanpes, shared with Mahakala, and differentiating them from Halszkaraptor. As such, appendicular disparity in the potentially sympatric halszkaraptorines suggest a reduced ecological overlap among these taxa, which may explain the co-occurrence of multiple species of this clade during the latest Cretaceous in what is now the Nemegt Basin.

BibTeX
@article{cau2018redescription,
    author = "Cau, Andrea and Madzia, Daniel",
    title = "Redescription and affinities of Hulsanpes perlei (Dinosauria, Theropoda) from the Upper Cretaceous of Mongolia",
    year = "2018",
    journal = "PeerJ",
    abstract = "Hulsanpes perlei is an enigmatic theropod dinosaur from the Baruungoyot Formation (?mid- to upper Campanian, Upper Cretaceous) of Mongolia. It was discovered in 1970, during the third Polish-Mongolian paleontological expedition to the Nemegt Basin. The taxon is known based on a partial braincase and an incomplete right hindlimb. However, the braincase fragment has never been described nor illustrated. We redescribe all elements that form the holotype of Hulsanpes and discuss the affinities of this taxon. The braincase fragment is interpreted as belonging to the inner ear region, and includes the floccular recess and part of the labyrinth. Hulsanpes perlei is confirmed as a valid taxon, diagnosed by a unique combination of metatarsal characters, including two autapomorphies. Historically, it represents the oldest record of the recently-established clade Halszkaraptorinae. Our findings identify subcursorial adaptations for Hulsanpes, shared with Mahakala, and differentiating them from Halszkaraptor. As such, appendicular disparity in the potentially sympatric halszkaraptorines suggest a reduced ecological overlap among these taxa, which may explain the co-occurrence of multiple species of this clade during the latest Cretaceous in what is now the Nemegt Basin.",
    url = "https://doi.org/10.7717/peerj.4868",
    doi = "10.7717/peerj.4868",
    openalex = "W2805542355",
    pages = "e4868",
    volume = "6",
    references = "doi101038415780a, doi101038nature03996, doi10108002724634199510011574, doi10108002724634200310010947, doi101111j10960031200800217x, doi101126science1144066, doi101127njgpm19821982440, doi1012067481, doi102475ajss321125417, doi105281zenodo16171435, openalexw1535663436, osmólska1982hulsanpes"
}

10. Gates, Terry A. and Tsogtbaatar, Khishigjav and Zanno, Lindsay E. and Chinzorig, Tsogtbaatar and Watabe, Mahito, 2018, A new iguanodontian (Dinosauria: Ornithopoda) from the Early Cretaceous of Mongolia: PeerJ: v. 6: p. e5300.

Abstract

We describe a new iguanodontian ornithopod, Choyrodon barsboldi gen. et sp. nov. from the Albian-aged Khuren Dukh Formation of Mongolia based on several partial skeletons interpreted to represent a subadult growth stage based on osteohistological features. This new taxon is diagnosed by many autapomorphies of the maxilla, nasal, lacrimal, opisthotic, predentary, and surangular. Choyrodon displays an unusual combination of traits, possessing an open antorbital fenestra (a primitive ornithopod trait) together with derived features such as a downturned dentary and enlarged narial fenestra. Histological imaging suggests that the type specimen of Choyrodon would have been a subadult at the time of death. Phylogenetic analysis of two different character matrices do not posit Choyrodon to be the sister taxon or to be more primitive than the iguanodontian Altirhinus kurzanovi, which is found in the same formation. The only resolved relationship of this new taxon is that it was hypothesized to be a sister-taxon with the North American species Eolambia caroljonesa. Though discovered in the same formation and Choyrodon being smaller-bodied than Altirhinus, it does not appear that the former species is an ontogimorph of the latter. Differences in morphology and results of the phylogenetic analyses support their distinction although more specimens of both species will allow better refinement of their uniqueness.

BibTeX
@article{gates2018a,
    author = "Gates, Terry A. and Tsogtbaatar, Khishigjav and Zanno, Lindsay E. and Chinzorig, Tsogtbaatar and Watabe, Mahito",
    title = "A new iguanodontian (Dinosauria: Ornithopoda) from the Early Cretaceous of Mongolia",
    year = "2018",
    journal = "PeerJ",
    abstract = "We describe a new iguanodontian ornithopod, Choyrodon barsboldi gen. et sp. nov. from the Albian-aged Khuren Dukh Formation of Mongolia based on several partial skeletons interpreted to represent a subadult growth stage based on osteohistological features. This new taxon is diagnosed by many autapomorphies of the maxilla, nasal, lacrimal, opisthotic, predentary, and surangular. Choyrodon displays an unusual combination of traits, possessing an open antorbital fenestra (a primitive ornithopod trait) together with derived features such as a downturned dentary and enlarged narial fenestra. Histological imaging suggests that the type specimen of Choyrodon would have been a subadult at the time of death. Phylogenetic analysis of two different character matrices do not posit Choyrodon to be the sister taxon or to be more primitive than the iguanodontian Altirhinus kurzanovi, which is found in the same formation. The only resolved relationship of this new taxon is that it was hypothesized to be a sister-taxon with the North American species Eolambia caroljonesa. Though discovered in the same formation and Choyrodon being smaller-bodied than Altirhinus, it does not appear that the former species is an ontogimorph of the latter. Differences in morphology and results of the phylogenetic analyses support their distinction although more specimens of both species will allow better refinement of their uniqueness.",
    url = "https://doi.org/10.7717/peerj.5300",
    doi = "10.7717/peerj.5300",
    openalex = "W2885937919",
    pages = "e5300",
    volume = "6",
    references = "doi1010079781489957405, doi101017s0094837300005820, doi101029sc005p0175, doi10108002724634199710011027, doi101242jeb00841, doi101525california97805202420980030023, doi1034191mp991, doi105860choice435902, openalexw3215057009, openalexw597685939"
}

11. Lee, Sungjin and Lee, Yuong-Nam and Chinsamy, Anusuya and Lü, Junchang and Barsbold, Rinchen and Tsogtbaatar, Khishigjav, 2019, A new baby oviraptorid dinosaur (Dinosauria: Theropoda) from the Upper Cretaceous Nemegt Formation of Mongolia: PLOS ONE: v. 14, no. 2: p. e0210867.

BibTeX
@article{lee2019a,
    author = "Lee, Sungjin and Lee, Yuong-Nam and Chinsamy, Anusuya and Lü, Junchang and Barsbold, Rinchen and Tsogtbaatar, Khishigjav",
    title = "A new baby oviraptorid dinosaur (Dinosauria: Theropoda) from the Upper Cretaceous Nemegt Formation of Mongolia",
    year = "2019",
    journal = "PLOS ONE",
    url = "https://doi.org/10.1371/journal.pone.0210867",
    doi = "10.1371/journal.pone.0210867",
    number = "2",
    openalex = "W2911778836",
    pages = "e0210867",
    volume = "14",
    references = "doi10103831635, doi101038378774a0, doi101073pnas0811087106, doi10108002724634199910011178, doi101111cla12160, doi101126science2665186779, doi101371journalpone0017114, doi105281zenodo16171435, doi105860choice353642, openalexw3215057009"
}

12. Tsogtbaatar, Khishigjav and Weishampel, David B. and Evans, David C. and Watabe, Mahito, 2019, A new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia): PLOS ONE: v. 14, no. 4: p. e0208480.

BibTeX
@article{tsogtbaatar2019a,
    author = "Tsogtbaatar, Khishigjav and Weishampel, David B. and Evans, David C. and Watabe, Mahito",
    title = "A new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia)",
    year = "2019",
    journal = "PLOS ONE",
    url = "https://doi.org/10.1371/journal.pone.0208480",
    doi = "10.1371/journal.pone.0208480",
    number = "4",
    openalex = "W2937612663",
    pages = "e0208480",
    volume = "14",
    references = "currie1993palaeontology, doi101007s0011401311075, doi1010160195667191900155, doi101017s1477201903001032, doi101017s1477201907002271, doi101080089129632012688589, doi101098rspl18870117, doi101111j10963642200900617x, doi101111zoj12193, doi101126science2725264986, doi101126science28454232137, doi101127njgpa210199841, doi101146annurevearth251435, doi105281zenodo16673433, openalexw2611511275, sereno1997the"
}

13. Averianov, Alexander O. and Sues, Hans‐Dieter, 2021, New material and diagnosis of a new taxon of alvarezsaurid (Dinosauria, Theropoda) from the Upper Cretaceous Bissekty Formation of Uzbekistan: Journal of Vertebrate Paleontology.

Abstract

A new alvarezsaurid theropod, Dzharaonyx eski, gen. et sp. nov., is documented by a variety of dissociated but well-preserved postcranial bones from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. It has opisthocoelous posterior dorsal vertebrae lacking infrapostzygapophyseal fossae. The posterior caudal vertebrae are procoelous and have a longitudinal canal within the neural arch. The humerus is intermediate in structure between those of the Argentinian alvarezsaurid Patagonykus and the Mongolian parvicursorine Mononykus in having a small internal tuberosity, similar-sized radial and ulnar condyles, a distally protruding radial condyle, and a greatly reduced entepicondyle. The ulna has a hypertrophied olecranon process, a globular carpal trochlea, and a facet for the aponeurosis tubercle of the radius. On the carpometacarpus, the articular surface of metacarpal III is in line with the distal articular joint of metacarpal II. On manual phalanx II-1, the flexor ridges are similarly developed and the dorsolateral process and the ‘hook-like’ ventrolateral process are absent. The collateral grooves on manual ungual phalanx II-2 are either partially open or completely enclosed to form ventral foramina. The pubis bears a preacetabular tubercle. The pes is arctometatarsalian. Metatarsal II forms a posterolateral flange. Pedal phalanx IV-1 has a protruding proximomedial process, a proximoventral notch, and asymmetrical distal condyles. The flexor tubercles on the pedal ungual phalanges are small. Phylogenetic analysis found Dzharaonyx within Parvicursorinae in a polytomy with other Asiatic taxa. Dzharaonyx is the most plesiomorphic and geologically oldest member of Parvicursorinae known to date.

BibTeX
@article{doi1010800272463420212036174,
    author = "Averianov, Alexander O. and Sues, Hans‐Dieter",
    title = "New material and diagnosis of a new taxon of alvarezsaurid (Dinosauria, Theropoda) from the Upper Cretaceous Bissekty Formation of Uzbekistan",
    year = "2021",
    journal = "Journal of Vertebrate Paleontology",
    abstract = "A new alvarezsaurid theropod, Dzharaonyx eski, gen. et sp. nov., is documented by a variety of dissociated but well-preserved postcranial bones from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. It has opisthocoelous posterior dorsal vertebrae lacking infrapostzygapophyseal fossae. The posterior caudal vertebrae are procoelous and have a longitudinal canal within the neural arch. The humerus is intermediate in structure between those of the Argentinian alvarezsaurid Patagonykus and the Mongolian parvicursorine Mononykus in having a small internal tuberosity, similar-sized radial and ulnar condyles, a distally protruding radial condyle, and a greatly reduced entepicondyle. The ulna has a hypertrophied olecranon process, a globular carpal trochlea, and a facet for the aponeurosis tubercle of the radius. On the carpometacarpus, the articular surface of metacarpal III is in line with the distal articular joint of metacarpal II. On manual phalanx II-1, the flexor ridges are similarly developed and the dorsolateral process and the ‘hook-like’ ventrolateral process are absent. The collateral grooves on manual ungual phalanx II-2 are either partially open or completely enclosed to form ventral foramina. The pubis bears a preacetabular tubercle. The pes is arctometatarsalian. Metatarsal II forms a posterolateral flange. Pedal phalanx IV-1 has a protruding proximomedial process, a proximoventral notch, and asymmetrical distal condyles. The flexor tubercles on the pedal ungual phalanges are small. Phylogenetic analysis found Dzharaonyx within Parvicursorinae in a polytomy with other Asiatic taxa. Dzharaonyx is the most plesiomorphic and geologically oldest member of Parvicursorinae known to date.",
    url = "https://doi.org/10.1080/02724634.2021.2036174",
    doi = "10.1080/02724634.2021.2036174",
    openalex = "W4221076854",
    references = "doi1010800891296320212000976"
}

14. Averianov, Alexander O. and Лопатин, А. В., 2021, The second taxon of alvarezsaurid theropod dinosaurs from the Late Cretaceous Khulsan locality in Gobi Desert, Mongolia: Historical Biology.

Abstract

Khulsanurus magnificus gen. et sp. nov., is described based on a partial skeleton, including cervical and caudal vertebrae, scapulocoracoids, humerus, and pubis from the Late Cretaceous (Campanian) Barungoyot Formation at Khulsan locality in Gobi Desert, Mongolia. The new taxon differs from other alvarezsaurids by a combination of characters that include cervicals lacking pleurocoels, carotid processes, and epipophyses, dorsoventrally thick and subtriangular in cross-section transverse process of the anterior caudals, prominent infrapostzygapophyseal fossa on the transverse process of anterior caudals, short and mostly anteriorly directed prezygapophyses of anterior caudals, neural arch of anterior caudal lacking the interzygapophyseal ridges and having a prominent dorsal depression around the anterior end of the neural spine. The new taxon retains a number of plesiomorphic traits: slightly convex posterior centrum condyle of anterior caudals, prominent anterior, and posterior bumps at the neurocentral junction in anterior caudals, neural arch of anterior caudals extending for the entire length of the centrum, and pubic foot and apron. Khulsanurus shares with Shuvuuia the deltopectoral crest of humerus that is continuous with the humeral head. The phylogenetic analysis placed Khulsanurus in the Parvicursorinae in a polytomy with Mononykus, Shuvuuia, Albinykus, and Xixianykus.

BibTeX
@article{doi1010800891296320212000976,
    author = "Averianov, Alexander O. and Лопатин, А. В.",
    title = "The second taxon of alvarezsaurid theropod dinosaurs from the Late Cretaceous Khulsan locality in Gobi Desert, Mongolia",
    year = "2021",
    journal = "Historical Biology",
    abstract = "Khulsanurus magnificus gen. et sp. nov., is described based on a partial skeleton, including cervical and caudal vertebrae, scapulocoracoids, humerus, and pubis from the Late Cretaceous (Campanian) Barungoyot Formation at Khulsan locality in Gobi Desert, Mongolia. The new taxon differs from other alvarezsaurids by a combination of characters that include cervicals lacking pleurocoels, carotid processes, and epipophyses, dorsoventrally thick and subtriangular in cross-section transverse process of the anterior caudals, prominent infrapostzygapophyseal fossa on the transverse process of anterior caudals, short and mostly anteriorly directed prezygapophyses of anterior caudals, neural arch of anterior caudal lacking the interzygapophyseal ridges and having a prominent dorsal depression around the anterior end of the neural spine. The new taxon retains a number of plesiomorphic traits: slightly convex posterior centrum condyle of anterior caudals, prominent anterior, and posterior bumps at the neurocentral junction in anterior caudals, neural arch of anterior caudals extending for the entire length of the centrum, and pubic foot and apron. Khulsanurus shares with Shuvuuia the deltopectoral crest of humerus that is continuous with the humeral head. The phylogenetic analysis placed Khulsanurus in the Parvicursorinae in a polytomy with Mononykus, Shuvuuia, Albinykus, and Xixianykus.",
    url = "https://doi.org/10.1080/08912963.2021.2000976",
    doi = "10.1080/08912963.2021.2000976",
    openalex = "W3213935139",
    references = "cau2018redescription, doi101016jcretres201111014, doi101016jcub201711007, doi101038362623a0, doi101080147720192013781067, doi101126science1182143, doi101127njgpm19821982440, doi1016660022336020030770822mbatho20co2, doi105281zenodo16171435, doi105860choice353642, openalexw1535663436, openalexw3215057009, osmólska1982hulsanpes"
}

15. Turner, Alan H. and Montanari, Shaena and Norell, Mark A., 2021, A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia: American Museum Novitates.

Abstract

A new dromaeosaurid dinosaur, Shri devi, from the Late Cretaceous deposit of the Barun Goyot Formation at Khulsan, Mongolia, is described here. The Barun Goyot Formation (herein referred to as the Barun Goyot) is stratigraphically intermediate between the overlying Nemegt Formation and the underlying Djadokhta Formation, where much of the dromaeosaurid diversity has been reported to date. Sediments of the Barun Goyot are typically considered Upper Campanian in age. Although dromaeosaurid remains have been noted to occur in the Barun Goyot for decades, descriptive and taxonomic work has never been completed for the material. The holotype specimen (IGM 100/980) consists of a partially articulated individual preserving the right hind limb; left tibiotarsus; pelvis; and adjacent cervical, dorsal, and caudal vertebrae. IGM 100/980 is referable to a clade with Velociraptor based on the presence of a distinct ambiens tubercle located proximally on the anterior face of the pubis, a well-developed anterior tuberosity proximally located on the ischium, and a rounded longitudinal ischial ridge. It is distinguishable from V. mongoliensis based on a weak fourth trochanter (shared with all other dromaeosaurids) and deep anterior pedicular fossae in the cervical vertebrae; epipophyses in the last four cervicals are not raised but are instead represented by rugose circular scars. A suite of axial and appendicular characters are diagnostic for the new species. New discoveries including Shri devi allow for an improved understanding of dromaeosaurid anatomy, as well as the temporal and regional variation of the dromaeosaurid fauna of Mongolia and Inner Mongolia (Nei Mongol Autonomous Region, China), during the Late Cretaceous.

BibTeX
@article{doi10120639651,
    author = "Turner, Alan H. and Montanari, Shaena and Norell, Mark A.",
    title = "A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia",
    year = "2021",
    journal = "American Museum Novitates",
    abstract = "A new dromaeosaurid dinosaur, Shri devi, from the Late Cretaceous deposit of the Barun Goyot Formation at Khulsan, Mongolia, is described here. The Barun Goyot Formation (herein referred to as the Barun Goyot) is stratigraphically intermediate between the overlying Nemegt Formation and the underlying Djadokhta Formation, where much of the dromaeosaurid diversity has been reported to date. Sediments of the Barun Goyot are typically considered Upper Campanian in age. Although dromaeosaurid remains have been noted to occur in the Barun Goyot for decades, descriptive and taxonomic work has never been completed for the material. The holotype specimen (IGM 100/980) consists of a partially articulated individual preserving the right hind limb; left tibiotarsus; pelvis; and adjacent cervical, dorsal, and caudal vertebrae. IGM 100/980 is referable to a clade with Velociraptor based on the presence of a distinct ambiens tubercle located proximally on the anterior face of the pubis, a well-developed anterior tuberosity proximally located on the ischium, and a rounded longitudinal ischial ridge. It is distinguishable from V. mongoliensis based on a weak fourth trochanter (shared with all other dromaeosaurids) and deep anterior pedicular fossae in the cervical vertebrae; epipophyses in the last four cervicals are not raised but are instead represented by rugose circular scars. A suite of axial and appendicular characters are diagnostic for the new species. New discoveries including Shri devi allow for an improved understanding of dromaeosaurid anatomy, as well as the temporal and regional variation of the dromaeosaurid fauna of Mongolia and Inner Mongolia (Nei Mongol Autonomous Region, China), during the Late Cretaceous.",
    url = "https://doi.org/10.1206/3965.1",
    doi = "10.1206/3965.1",
    openalex = "W3118766077",
    references = "doi101016s0748300703000604, doi10103835047056, doi10108002724634199910011178, doi101086273307, doi101111cla12160, doi101111j109600312003tb00376x, doi101126science1144066, doi101127njgpm19821982440, doi1012067481, doi101525california97805202420980010001, doi107717peerj4558, openalexw617951419, osmólska1982hulsanpes"
}

16. Napoli, James G. and Ruebenstahl, Alexander and Bhullar, Bhart‐Anjan S. and Turner, Alan H. and Norell, Mark A., 2021, A New Dromaeosaurid (Dinosauria: Coelurosauria) from Khulsan, Central Mongolia: American Museum Novitates.

Abstract

Dromaeosaurid theropods represent a rare but important clade of nonavialan dinosaurs. Their close evolutionary relationship to modern birds has placed them at the center of paleontological research for the last several decades. Herein we describe a new species of dromaeosaurid—Kuru kulla, gen. et sp. nov.—based on a partial skeleton from the Late Cretaceous Khulsan locality (Barun Goyot Formation) of Mongolia. This species is diagnosed by several autapomorphies within Dromaeosauridae, including a sharp groove anterior and ventral to the narial fossa on the premaxilla, a posterolaterally directed hornlet on the posterodorsal process of the lacrimal, a deep surangular bearing two surangular foramina, and anteriorly displaced pleurocoels on the dorsal centra. The taxon is further characterized by a unique combination of characters, including a mediolaterally narrow metatarsal II, serrations on both carinae of the dentary teeth, hyposphenes that are widely separated but joined by a web of bone, and a lacrimal with a poorly developed boss on its lateral surface. Phylogenetic analysis finds Kuru kulla to be the sister taxon of Adasaurus mongoliensis, from the slightly later Nemegt Formation, with which it is united by three synapomorphies: a posterior surangular foramen that is ∼30% the depth of the surangular, absence of a fourth trochanter of the femur, and thoracic centra that are markedly longer than their midpoint widths. The recognition of this taxon has important implications for common assumptions of Mesozoic terrestrial ecosystem structure and adds new data to a recently recognized pattern in dromaeosaurid faunal composition among Late Cretaceous localities in Mongolia and Inner Mongolia (Nei Mongol Autonomous Region, China).

BibTeX
@article{doi10120639821,
    author = "Napoli, James G. and Ruebenstahl, Alexander and Bhullar, Bhart‐Anjan S. and Turner, Alan H. and Norell, Mark A.",
    title = "A New Dromaeosaurid (Dinosauria: Coelurosauria) from Khulsan, Central Mongolia",
    year = "2021",
    journal = "American Museum Novitates",
    abstract = "Dromaeosaurid theropods represent a rare but important clade of nonavialan dinosaurs. Their close evolutionary relationship to modern birds has placed them at the center of paleontological research for the last several decades. Herein we describe a new species of dromaeosaurid—Kuru kulla, gen. et sp. nov.—based on a partial skeleton from the Late Cretaceous Khulsan locality (Barun Goyot Formation) of Mongolia. This species is diagnosed by several autapomorphies within Dromaeosauridae, including a sharp groove anterior and ventral to the narial fossa on the premaxilla, a posterolaterally directed hornlet on the posterodorsal process of the lacrimal, a deep surangular bearing two surangular foramina, and anteriorly displaced pleurocoels on the dorsal centra. The taxon is further characterized by a unique combination of characters, including a mediolaterally narrow metatarsal II, serrations on both carinae of the dentary teeth, hyposphenes that are widely separated but joined by a web of bone, and a lacrimal with a poorly developed boss on its lateral surface. Phylogenetic analysis finds Kuru kulla to be the sister taxon of Adasaurus mongoliensis, from the slightly later Nemegt Formation, with which it is united by three synapomorphies: a posterior surangular foramen that is ∼30\% the depth of the surangular, absence of a fourth trochanter of the femur, and thoracic centra that are markedly longer than their midpoint widths. The recognition of this taxon has important implications for common assumptions of Mesozoic terrestrial ecosystem structure and adds new data to a recently recognized pattern in dromaeosaurid faunal composition among Late Cretaceous localities in Mongolia and Inner Mongolia (Nei Mongol Autonomous Region, China).",
    url = "https://doi.org/10.1206/3982.1",
    doi = "10.1206/3982.1",
    openalex = "W3212801756",
    references = "doi101002ar24241, doi10120639651"
}

17. Funston, Gregory F. and Currie, Philip J. and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav, 2021, A partial oviraptorosaur skeleton suggests low caenagnathid diversity in the Late Cretaceous Nemegt Formation of Mongolia: PLoS ONE.

Abstract

The Nemegt Formation of the Gobi Desert of Mongolia has produced one of the most abundant and diverse oviraptorosaur records globally. However, the caenagnathid component of this fauna remains poorly known. Two caenagnathid taxa are currently recognized from the Nemegt Formation: Elmisaurus rarus and Nomingia gobiensis. Because these taxa are known from mostly non-overlapping material, there are concerns that they could represent the same animal. A partial, weathered caenagnathid skeleton discovered adjacent to the holotype quarry of Nomingia gobiensis is referable to Elmisaurus rarus, revealing more of the morphology of the cranium, mandible, pectoral girdle, and pubis. Despite metatarsals clearly exhibiting autapomorphies of Elmisaurus rarus, overlapping elements are identical to those of Nomingia gobiensis, and add to a growing body of evidence that these taxa represent a single morphotype. In the absence of any positive evidence for two caenagnathid taxa in the Nemegt Formation, Nomingia gobiensis is best regarded as a junior synonym of Elmisaurus rarus. Low caenagnathid diversity in the Nemegt Formation may reflect broader coexistence patterns with other oviraptorosaur families, particularly oviraptorids. In contrast to North America, competition with the exceptionally diverse oviraptorids may have restricted caenagnathids to marginal roles in Late Cretaceous Asian ecosystems.

BibTeX
@article{doi101371journalpone0254564,
    author = "Funston, Gregory F. and Currie, Philip J. and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav",
    title = "A partial oviraptorosaur skeleton suggests low caenagnathid diversity in the Late Cretaceous Nemegt Formation of Mongolia",
    year = "2021",
    journal = "PLoS ONE",
    abstract = "The Nemegt Formation of the Gobi Desert of Mongolia has produced one of the most abundant and diverse oviraptorosaur records globally. However, the caenagnathid component of this fauna remains poorly known. Two caenagnathid taxa are currently recognized from the Nemegt Formation: Elmisaurus rarus and Nomingia gobiensis. Because these taxa are known from mostly non-overlapping material, there are concerns that they could represent the same animal. A partial, weathered caenagnathid skeleton discovered adjacent to the holotype quarry of Nomingia gobiensis is referable to Elmisaurus rarus, revealing more of the morphology of the cranium, mandible, pectoral girdle, and pubis. Despite metatarsals clearly exhibiting autapomorphies of Elmisaurus rarus, overlapping elements are identical to those of Nomingia gobiensis, and add to a growing body of evidence that these taxa represent a single morphotype. In the absence of any positive evidence for two caenagnathid taxa in the Nemegt Formation, Nomingia gobiensis is best regarded as a junior synonym of Elmisaurus rarus. Low caenagnathid diversity in the Nemegt Formation may reflect broader coexistence patterns with other oviraptorosaur families, particularly oviraptorids. In contrast to North America, competition with the exceptionally diverse oviraptorids may have restricted caenagnathids to marginal roles in Late Cretaceous Asian ecosystems.",
    url = "https://doi.org/10.1371/journal.pone.0254564",
    doi = "10.1371/journal.pone.0254564",
    openalex = "W3179240167",
    references = "doi101016jcretres2020104521, doi1018435vamp29362, lee2019a"
}

18. Son, Minyoung and Lee, Yuong-Nam and Zorigt, Badamkhatan and Kobayashi, Yoshitsugu and Park, Jin-Young and Lee, Sungjin and Kim, Su-Hwan and Lee, Kang Young, 2022, A new juvenile Yamaceratops (Dinosauria, Ceratopsia) from the Javkhlant Formation (Upper Cretaceous) of Mongolia: PeerJ: v. 10: p. e13176.

Abstract

Here we report a new articulated skeleton of Yamaceratops dorngobiensis (MPC-D 100/553) from the Khugenetjavkhlant locality at the Shine Us Khudag (Javkhlant Formation,?Santonian-Campanian) of the eastern Gobi Desert, Mongolia, which represents the first substantially complete skeleton and the first juvenile individual of this taxon. The specimen includes a nearly complete cranium and large portions of the vertebral column and appendicular skeleton. Its skull is about 2/3 the size of the holotype specimen, based on mandibular length. Its juvenile ontogenetic stage is confirmed by multiple indicators of skeletal and morphological immaturity known in ceratopsians, such as the long-grained surface texture on the long bones, the smooth external surface on the postorbital, open neurocentral sutures of all caudal vertebrae, a large orbit relative to the postorbital and jugal, the low angle of the lacrimal ventral ramus relative to the maxillary teeth row, narrow frontal, and straight ventral edge of the dentary. Osteohistological analysis of MPC-D 100/553 recovered three lines of arrested growth, implying around 3 years of age when it died, and verified this specimen’s immature ontogenetic stage. The specimen adds a new autapomorphy of Yamaceratops, the anteroventral margin of the fungiform dorsal end of the lacrimal being excluded from the antorbital fossa. Furthermore, it shows a unique combination of diagnostic features of some other basal neoceratopsians: the ventrally hooked rostral bone as in Aquilops americanus and very tall middle caudal neural spines about or more than four times as high as the centrum as in Koreaceratops hwaseongensis, Montanoceratops cerorhynchus, and Protoceratops andrewsi. The jugal with the subtemporal ramus deeper than the suborbital ramus as in the holotype specimen is also shared with A. americanus, Liaoceratops yanzigouensis, and juvenile P. andrewsi. Adding 38 new scorings into the recent comprehensive data matrix of basal Neoceratopsia and taking into account the ontogenetically variable characters recovered Y. dorngobiensis as the sister taxon to Euceratopsia (Leptoceratopsidae plus Coronosauria). A second phylogenetic analysis with another matrix for Ceratopsia also supported this position. The new phylogenetic position of Y. dorngobiensis is important in ceratopsian evolution, as this taxon represents one of the basalmost neoceratopsians with a broad, thin frill and hyper-elongated middle caudal neural spines while still being bipedal.

BibTeX
@article{son2022a,
    author = "Son, Minyoung and Lee, Yuong-Nam and Zorigt, Badamkhatan and Kobayashi, Yoshitsugu and Park, Jin-Young and Lee, Sungjin and Kim, Su-Hwan and Lee, Kang Young",
    title = "A new juvenile Yamaceratops (Dinosauria, Ceratopsia) from the Javkhlant Formation (Upper Cretaceous) of Mongolia",
    year = "2022",
    journal = "PeerJ",
    abstract = "Here we report a new articulated skeleton of Yamaceratops dorngobiensis (MPC-D 100/553) from the Khugenetjavkhlant locality at the Shine Us Khudag (Javkhlant Formation,?Santonian-Campanian) of the eastern Gobi Desert, Mongolia, which represents the first substantially complete skeleton and the first juvenile individual of this taxon. The specimen includes a nearly complete cranium and large portions of the vertebral column and appendicular skeleton. Its skull is about 2/3 the size of the holotype specimen, based on mandibular length. Its juvenile ontogenetic stage is confirmed by multiple indicators of skeletal and morphological immaturity known in ceratopsians, such as the long-grained surface texture on the long bones, the smooth external surface on the postorbital, open neurocentral sutures of all caudal vertebrae, a large orbit relative to the postorbital and jugal, the low angle of the lacrimal ventral ramus relative to the maxillary teeth row, narrow frontal, and straight ventral edge of the dentary. Osteohistological analysis of MPC-D 100/553 recovered three lines of arrested growth, implying around 3 years of age when it died, and verified this specimen’s immature ontogenetic stage. The specimen adds a new autapomorphy of Yamaceratops, the anteroventral margin of the fungiform dorsal end of the lacrimal being excluded from the antorbital fossa. Furthermore, it shows a unique combination of diagnostic features of some other basal neoceratopsians: the ventrally hooked rostral bone as in Aquilops americanus and very tall middle caudal neural spines about or more than four times as high as the centrum as in Koreaceratops hwaseongensis, Montanoceratops cerorhynchus, and Protoceratops andrewsi. The jugal with the subtemporal ramus deeper than the suborbital ramus as in the holotype specimen is also shared with A. americanus, Liaoceratops yanzigouensis, and juvenile P. andrewsi. Adding 38 new scorings into the recent comprehensive data matrix of basal Neoceratopsia and taking into account the ontogenetically variable characters recovered Y. dorngobiensis as the sister taxon to Euceratopsia (Leptoceratopsidae plus Coronosauria). A second phylogenetic analysis with another matrix for Ceratopsia also supported this position. The new phylogenetic position of Y. dorngobiensis is important in ceratopsian evolution, as this taxon represents one of the basalmost neoceratopsians with a broad, thin frill and hyper-elongated middle caudal neural spines while still being bipedal.",
    url = "https://doi.org/10.7717/peerj.13176",
    doi = "10.7717/peerj.13176",
    openalex = "W4226079443",
    pages = "e13176",
    volume = "10",
    references = "allison1988the, doi101002ar24094, doi1010160305440381900066, doi101017s0094837300005820, doi101038nmeth2089, doi10108002724634199610011283, doi1010800272463420181509866, doi1011112041210x12226, doi101111cla12160, doi101111pala12142, doi101134s0031030120120047, openalexw1535663436, openalexw3215057009"
}

19. Li, Yang and Li, Xiaobo and Chen, Jun and Reisz, Robert R., 2023, First discovery of large-bodied dromaeosaurid fossil materials (Dinosauria: Theropoda) from the Upper Cretaceous Quantou Formation, Songliao Basin, Northeast China: Cretaceous Research.

BibTeX
@article{doi101016jcretres2023105711,
    author = "Li, Yang and Li, Xiaobo and Chen, Jun and Reisz, Robert R.",
    title = "First discovery of large-bodied dromaeosaurid fossil materials (Dinosauria: Theropoda) from the Upper Cretaceous Quantou Formation, Songliao Basin, Northeast China",
    year = "2023",
    journal = "Cretaceous Research",
    url = "https://doi.org/10.1016/j.cretres.2023.105711",
    doi = "10.1016/j.cretres.2023.105711",
    openalex = "W4386840802",
    references = "doi101002spp21375, doi10120639651, doi101371journalpone0205206"
}

20. Kubo, Kohta and Kobayashi, Yoshitsugu and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav, 2023, A new alvarezsaurid dinosaur (Theropoda, Alvarezsauria) from the Upper Cretaceous Baruungoyot Formation of Mongolia provides insights for bird-like sleeping behavior in non-avian dinosaurs.: PloS one.

Abstract

Alvarezsauria is a group of early-branching maniraptoran theropods that are distributed globally from the Late Jurassic to the latest Cretaceous. Despite recent increases in the fossil record of this group, the scarcity of complete specimens still restricts interpreting their detailed anatomy, ecology, and evolution. Here, we report a new taxon of derived alvarezsaur, Jaculinykus yaruui gen. et sp. nov., from the Late Cretaceous of Mongolia, which represents a nearly complete and articulated skeleton. Our phylogenetic analysis reveals that Jaculinykus belongs to the sub-clade of Alvarezsauridae, Parvicursorinae, and forms a mononphyletic group with Mononykus and Shuvuuia. Its well-preserved manus has only two fingers, composed of a hypertrophied digit I and greatly reduced digit II, which implies an intermediate condition between the tridactyl manus of Shuvuuia and monodactyl manus of Linhenykus. This highlights a previously unrecognized variation in specialization of alvarezsaurid manus. Notably, the preserved posture of the specimen exhibits a stereotypical avian-like sleeping position seen in the troodontids Mei and Sinornithoides. Evidence of this behavior in the alvarezsaur Jaculinykus suggests that stereotypically avian sleeping postures are a maniraptoran synapomorphy, providing more evidence of bird-like traits being distributed broadly among avian ancestors.

BibTeX
@article{doi101371journalpone0293801,
    author = "Kubo, Kohta and Kobayashi, Yoshitsugu and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav",
    title = "A new alvarezsaurid dinosaur (Theropoda, Alvarezsauria) from the Upper Cretaceous Baruungoyot Formation of Mongolia provides insights for bird-like sleeping behavior in non-avian dinosaurs.",
    year = "2023",
    journal = "PloS one",
    abstract = "Alvarezsauria is a group of early-branching maniraptoran theropods that are distributed globally from the Late Jurassic to the latest Cretaceous. Despite recent increases in the fossil record of this group, the scarcity of complete specimens still restricts interpreting their detailed anatomy, ecology, and evolution. Here, we report a new taxon of derived alvarezsaur, Jaculinykus yaruui gen. et sp. nov., from the Late Cretaceous of Mongolia, which represents a nearly complete and articulated skeleton. Our phylogenetic analysis reveals that Jaculinykus belongs to the sub-clade of Alvarezsauridae, Parvicursorinae, and forms a mononphyletic group with Mononykus and Shuvuuia. Its well-preserved manus has only two fingers, composed of a hypertrophied digit I and greatly reduced digit II, which implies an intermediate condition between the tridactyl manus of Shuvuuia and monodactyl manus of Linhenykus. This highlights a previously unrecognized variation in specialization of alvarezsaurid manus. Notably, the preserved posture of the specimen exhibits a stereotypical avian-like sleeping position seen in the troodontids Mei and Sinornithoides. Evidence of this behavior in the alvarezsaur Jaculinykus suggests that stereotypically avian sleeping postures are a maniraptoran synapomorphy, providing more evidence of bird-like traits being distributed broadly among avian ancestors.",
    url = "https://pmc.ncbi.nlm.nih.gov/articles/PMC10651048/",
    doi = "10.1371/journal.pone.0293801",
    pmcid = "PMC10651048",
    pmid = "37967055"
}

21. Lee, Sungjin and Lee, Yuong-Nam and Park, Jin-Young and Kim, Su-Hwan and Badamkhatan, Zorigt and Idersaikhan, Damdinsuren and Tsogtbaatar, Khishigjav, 2023, The first troodontid (Dinosauria: Theropoda) from the Upper Cretaceous Baruungoyot Formation of Mongolia: Journal of Vertebrate Paleontology: v. 43, no. 6.

BibTeX
@article{lee2023the,
    author = "Lee, Sungjin and Lee, Yuong-Nam and Park, Jin-Young and Kim, Su-Hwan and Badamkhatan, Zorigt and Idersaikhan, Damdinsuren and Tsogtbaatar, Khishigjav",
    title = "The first troodontid (Dinosauria: Theropoda) from the Upper Cretaceous Baruungoyot Formation of Mongolia",
    year = "2023",
    journal = "Journal of Vertebrate Paleontology",
    url = "https://doi.org/10.1080/02724634.2024.2364746",
    doi = "10.1080/02724634.2024.2364746",
    number = "6",
    volume = "43"
}

22. Tucker, Ryan T. and King, M. Ryan and Delgerzaya, Puntsag and Freimuth, William J. and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav and Takasaki, Ryuji and Saneyoshi, Mototaka and Chiarenza, Alfio Alessandro and Zanno, Lindsay E., 2025, Protracted intercontinental aridification preserved within the early Late Cretaceous strata of the Eastern Gobi Basin, Mongolia: Sedimentology.

Abstract

ABSTRACT Mongolia's Eastern and Western Gobi Basins preserve a globally significant record of Cretaceous terrestrial vertebrates, yet their biostratigraphic correlations are complicated by a complex geological history. The Eastern Gobi Basin, a northeast‐southwest trending fault‐bounded rift system, includes several minor sub‐basins with distinct sedimentary sequences, notably the Upper Cretaceous Bayanshiree Formation. This formation hosts key localities of iconic taxa, including Segnosaurus, Erlikosaurus, Duonychus, Garudimimus, Amtocephale, Gobihadros and Adocus amtgai, yet previous correlations throughout the area lacked precision. Our sedimentological and stratigraphic campaigns (2022 to 2024) in the Bayanshiree Formation and overlying red‐bed sequences at Baishin Tsav refined these biostratigraphic and palaeoenvironmental frameworks. Field surveys at local (Baishin Tsav) and basin‐wide scales (Unegt and Zuunbayan sub‐basins) revealed a significant erosional unconformity dividing the formation into two distinct palaeoenvironmental stages: lower expansive erg (aeolian dunes) and upper mature fluvial floodplains. Additionally, this study redefines the previously misidentified red‐bed sequences above the Bayanshiree formation as the Javkhlant Formation (formerly Djadokhta or Baruungoyot formations). Taphonomic analyses indicate fossil assemblages were extensively reworked, demonstrating significant time‐averaging. This study provides evidence for climatic shifts through three successive environmental phases (erg, floodplain, alternating palaeosols) within the Bayanshiree and Javkhlant formations. These findings suggest widespread aridification in eastern Asia initiated during the Cenomanian–Turonian transition, contemporaneous with a subtropical high‐pressure shift and onset of the Cretaceous Thermal Maximum, challenging prior assumptions that placed this event later in the Late Cretaceous. This palaeoclimatic interpretation aligns with global records of past hyperthermal events and significantly refines the temporal context for interpreting regional palaeobiodiversity patterns.

BibTeX
@article{doi101111sed70054,
    author = "Tucker, Ryan T. and King, M. Ryan and Delgerzaya, Puntsag and Freimuth, William J. and Chinzorig, Tsogtbaatar and Tsogtbaatar, Khishigjav and Takasaki, Ryuji and Saneyoshi, Mototaka and Chiarenza, Alfio Alessandro and Zanno, Lindsay E.",
    title = "Protracted intercontinental aridification preserved within the early Late Cretaceous strata of the Eastern Gobi Basin, Mongolia",
    year = "2025",
    journal = "Sedimentology",
    abstract = "ABSTRACT Mongolia's Eastern and Western Gobi Basins preserve a globally significant record of Cretaceous terrestrial vertebrates, yet their biostratigraphic correlations are complicated by a complex geological history. The Eastern Gobi Basin, a northeast‐southwest trending fault‐bounded rift system, includes several minor sub‐basins with distinct sedimentary sequences, notably the Upper Cretaceous Bayanshiree Formation. This formation hosts key localities of iconic taxa, including Segnosaurus, Erlikosaurus, Duonychus, Garudimimus, Amtocephale, Gobihadros and Adocus amtgai, yet previous correlations throughout the area lacked precision. Our sedimentological and stratigraphic campaigns (2022 to 2024) in the Bayanshiree Formation and overlying red‐bed sequences at Baishin Tsav refined these biostratigraphic and palaeoenvironmental frameworks. Field surveys at local (Baishin Tsav) and basin‐wide scales (Unegt and Zuunbayan sub‐basins) revealed a significant erosional unconformity dividing the formation into two distinct palaeoenvironmental stages: lower expansive erg (aeolian dunes) and upper mature fluvial floodplains. Additionally, this study redefines the previously misidentified red‐bed sequences above the Bayanshiree formation as the Javkhlant Formation (formerly Djadokhta or Baruungoyot formations). Taphonomic analyses indicate fossil assemblages were extensively reworked, demonstrating significant time‐averaging. This study provides evidence for climatic shifts through three successive environmental phases (erg, floodplain, alternating palaeosols) within the Bayanshiree and Javkhlant formations. These findings suggest widespread aridification in eastern Asia initiated during the Cenomanian–Turonian transition, contemporaneous with a subtropical high‐pressure shift and onset of the Cretaceous Thermal Maximum, challenging prior assumptions that placed this event later in the Late Cretaceous. This palaeoclimatic interpretation aligns with global records of past hyperthermal events and significantly refines the temporal context for interpreting regional palaeobiodiversity patterns.",
    url = "https://doi.org/10.1111/sed.70054",
    doi = "10.1111/sed.70054",
    openalex = "W4416344771",
    references = "doi101016jearscirev2023104418, doi101016jsedgeo200512005, doi107717peerj11290, son2022a"
}