@article{doi101038298425a0,
    author = "Grafen, Alan",
    title = "How not to measure inclusive fitness",
    year = "1982",
    journal = "Nature",
    url = "https://doi.org/10.1038/298425a0",
    doi = "10.1038/298425a0",
    openalex = "W2042966014"
}

@article{doi1010160040580988900391,
    author = "Taylor, Peter",
    title = "Inclusive fitness models with two sexes",
    year = "1988",
    journal = "Theoretical Population Biology",
    url = "https://doi.org/10.1016/0040-5809(88)90039-1",
    doi = "10.1016/0040-5809(88)90039-1",
    openalex = "W2005795075"
}

@article{doi101016s0022519388800353,
    author = "Taylor, Peter",
    title = "An inclusive fitness model for dispersal of offspring",
    year = "1988",
    journal = "Journal of Theoretical Biology",
    url = "https://doi.org/10.1016/s0022-5193(88)80035-3",
    doi = "10.1016/s0022-5193(88)80035-3",
    openalex = "W2040715891"
}

@article{doi101007bf02270971,
    author = "Taylor, Peter",
    title = "Altruism in viscous populations — an inclusive fitness model",
    year = "1992",
    journal = "Evolutionary Ecology",
    url = "https://doi.org/10.1007/bf02270971",
    doi = "10.1007/bf02270971",
    openalex = "W2035440806"
}

@article{doi101086285343,
    author = "Queller, David C.",
    title = "Quantitative Genetics, Inclusive Fitness, and Group Selection",
    year = "1992",
    journal = "The American Naturalist",
    abstract = "Inclusive-fitness models have been criticized because they give incorrect results for cases in which fitness components interact nonadditively. However, this failure is not due to anything intrinsic to the inclusive-fitness viewpoint. It stems from an essentially quantitative genetic feature of the model, an attempt to separate fitness terms from genetic terms. A general rule is provided for determining when such a separation is justified. This rule is used to show how Price's covariance equation is related to standard quantitative genetic results and to derive quantitative genetic equations for inclusive fitness and group selection. It also shows that the group-selection model is no more general than the inclusive-fitness viewpoint. These models serve a role that is different from, but not inferior to, population-genetics models. Although they are less exact under some conditions, like quantitative genetic models in general, they provide us with measurable parameters.",
    url = "https://doi.org/10.1086/285343",
    doi = "10.1086/285343",
    openalex = "W2090834726"
}

@article{doi10103700223514675773,
    author = "Burnstein, Eugene and Crandall, Christian S. and Kitayama, Shinobu",
    title = "Some neo-Darwinian decision rules for altruism: Weighing cues for inclusive fitness as a function of the biological importance of the decision.",
    year = "1994",
    journal = "Journal of Personality and Social Psychology",
    abstract = "A neo-Darwinian heuristic for decisions involving altruism is'hypothesized in this article. Following W. Hamilton's (1964) analysis of inclusive fitness, the assumption of this study is that tendencies to help another person are selected against, except when the donor and recipient are related and share genes underlying these tendencies. An important social psychological implication of Hamilton's formulation is that in group-living individuals (a) natural selection favors those who are prone to help others as a function of the latters' relatedness, potential fecundity, or other features indicating a recipient's capacity to enhance the donors' inclusive fitness, and (b) this effect is especially strong when help is biologically significant (e.g., the recipient will not survive otherwise). Such a heuristic is demonstrated in several studies involving hypothetical decisions to help: In life-or-death situations, people chose to aid close kin over distant kin, the young over the old, the healthy over the sick, the wealthy over the poor, and the premenopausal woman over the postmenopausal woman; whereas when it is a matter of an everyday favor, they gave less weight to kinship and opted to help either the very young or the very old over those of intermediate age, the sick over the healthy, and the poor over the wealthy.",
    url = "https://doi.org/10.1037/0022-3514.67.5.773",
    doi = "10.1037/0022-3514.67.5.773",
    openalex = "W1994205665"
}

@article{doi101007bf02409753,
    author = "Taylor, Peter",
    title = "Inclusive fitness arguments in genetic models of behaviour",
    year = "1996",
    journal = "Journal of Mathematical Biology",
    url = "https://doi.org/10.1007/bf02409753",
    doi = "10.1007/bf02409753",
    openalex = "W2090682894"
}

@article{doi101016jjtbi200506009,
    author = "Grafen, Alan",
    title = "Optimization of inclusive fitness",
    year = "2005",
    journal = "Journal of Theoretical Biology",
    url = "https://doi.org/10.1016/j.jtbi.2005.06.009",
    doi = "10.1016/j.jtbi.2005.06.009",
    openalex = "W2062827139"
}

@article{doi101038nature09831,
    author = "Abbot, Patrick and Abe, Jun and Alcock, John and Alizon, Samuel and Alpedrinha, João and Andersson, Malte and André, Jean‐Baptiste and van Baalen, Minus and Balloux, François and Balshine, Sigal and Barton, Nick and Beukeboom, Leo W. and Biernaskie, Jay M. and Bilde, Trine and Borgia, Gerald and Breed, Michael D. and Brown, Sam P. and Bshary, Redouan and Buckling, Angus and Burley, Nancy Tyler and Burton-Chellew, Maxwell N. and Cant, Michael A. and Chapuisat, Michel and Charnov, Eric L. and Clutton‐Brock, Tim and Cockburn, Andrew and Cole, Blaine J. and Colegrave, Nick and Cosmides, Leda and Couzin, Iain D. and Coyne, Jerry A. and Creel, Scott and Crespi, Bernard J. and Curry, Robert L. and Dall, Sasha R. X. and Day, Troy and Dickinson, Janis L. and Dugatkin, Lee Alan and Mouden, Claire El and Emlen, Stephen T. and Evans, Jay D. and Ferrière, Régis and Field, Jeremy and Foitzik, Susanne and Foster, Kevin R. and Foster, William A. and Fox, Charles W. and Gadau, Juergen and Gandon, Sylvain and Gardner, Andy and Gardner, M. and Getty, Thomas and Goodisman, Michael A. D. and Grafen, Alan and Grosberg, Rick and Grozinger, Christina M. and Gouyon, Pierre-Henri and Gwynne, Darryl and Harvey, Paul and Hatchwell, Ben J. and Heınze, Jürgen and Helanterä, Heikki and Helms, Ken R. and Hill, Kim and Jiricny, Natalie and Johnstone, Rufus A. and Kacelnik, Alex and Kiers, E. Toby and Kokko, Hanna and Komdeur, Jan and Korb, Judith and Kronauer, Daniel J. C. and Kümmerli, Rolf and Lehmann, Laurent and Linksvayer, Timothy A. and Lion, Sébastien and Lyon, Bruce E. and Marshall, James A. R. and McElreath, Richard and Michalakis, Yannis and Michod, Richard E. and Mock, Douglas W. and Monnin, Thibaud and Montgomerie, Robert and Moore, Allen J. and Mueller, Ulrich G. and Noë, Ronald and Okasha, Samir and Pamilo, Pekka and Parker, Geoff A. and Pedersen, Jes Søe and Pen, Ido and Pfennig, David W. and Queller, David C. and Rankin, Daniel J. and Reece, Sarah E. and Reeve, H. Kern and Reuter, Max and Roberts, Gilbert and Robson, Simon K. A.",
    title = "Inclusive fitness theory and eusociality",
    year = "2011",
    journal = "Nature",
    url = "https://doi.org/10.1038/nature09831",
    doi = "10.1038/nature09831",
    openalex = "W2151939613"
}

@article{doi101038nature09834,
    author = "Ferrière, Régis and Michod, Richard E.",
    title = "Inclusive fitness in evolution",
    year = "2011",
    journal = "Nature",
    url = "https://doi.org/10.1038/nature09834",
    doi = "10.1038/nature09834",
    openalex = "W2154447920"
}

@article{doi101016jcub201305031,
    author = "West, Stuart A. and Gardner, Andy",
    title = "Adaptation and Inclusive Fitness",
    year = "2013",
    journal = "Current Biology",
    url = "https://doi.org/10.1016/j.cub.2013.05.031",
    doi = "10.1016/j.cub.2013.05.031",
    openalex = "W2169140583"
}

@article{doi101073pnas1317588110,
    author = "Allen, Benjamin and Nowak, Martin A. and Wilson, Edward O.",
    title = "Limitations of inclusive fitness",
    year = "2013",
    journal = "Proceedings of the National Academy of Sciences",
    abstract = "Until recently, inclusive fitness has been widely accepted as a general method to explain the evolution of social behavior. Affirming and expanding earlier criticism, we demonstrate that inclusive fitness is instead a limited concept, which exists only for a small subset of evolutionary processes. Inclusive fitness assumes that personal fitness is the sum of additive components caused by individual actions. This assumption does not hold for the majority of evolutionary processes or scenarios. To sidestep this limitation, inclusive fitness theorists have proposed a method using linear regression. On the basis of this method, it is claimed that inclusive fitness theory (i) predicts the direction of allele frequency changes, (ii) reveals the reasons for these changes, (iii) is as general as natural selection, and (iv) provides a universal design principle for evolution. In this paper we evaluate these claims, and show that all of them are unfounded. If the objective is to analyze whether mutations that modify social behavior are favored or opposed by natural selection, then no aspect of inclusive fitness theory is needed.",
    url = "https://doi.org/10.1073/pnas.1317588110",
    doi = "10.1073/pnas.1317588110",
    openalex = "W2080570670"
}